
The cornulitids
The exact
biological affinities of cornulitids are unknown
(Fisher, 1962; Morris and Rollins 1971; Richards 1974; Gnoli
1992). Four genera Cornulites Schlotheim, 1820, Conchicolites
Nicholson, 1872a, Cornulitella Nicholson,
1872b, Kolihaia Prantl,
1944, have been assigned to Cornulitidae (Fisher
1962). They have been affiliated to coelenterates (stromatoporoids,
calcareous hydroids), fusulines, bryozoans, annelids and mollusks (Schlotheim
1820, Nicholson 1872a, Nicholson 1872b, Moore et al. 1952; La Rocque and Marple
1955; Fisher 1962; Bilnd 1972; Dzik 1991; Kriz et al. 2001).
Dzik (1991) erected a new family Cornulitozoidae
within the order Cornulitida (Bouãek
1964), containing two new genera, Cornulitozoon
and Opatozoon, from Upper Ordovician
and Silurian of Poland. Cornulitids range from Middle
Ordovician to
Mississippian (Fisher 1962; Richards 1974) and vary morphologically from single
tubes to clusters of tubes cemented to each other and radiating outward in a
fan-like manner. Cornulites shares most of
characters with the lophoporates and was probably
linked to tentaculitids
and microconchids
(Vinn and Mutvei, 2005).
Shell structures of Cornulites
Longitudinal section of
the vesicular wall structure in Cornulites cf.
proprius
Cross-section of the
laminar shell with the pseudopunctae in C. cf.
scalariformis
Juvenile specimens
with the embryonic shells preserved in C. cf. flexuosus
Shell structures of Septalites
Longitudinal
section of Septalites septatus
Vinn, 2005 shells with multiple septa; Silurian of
Simple
prismatic structure and lamellar structure in the Septalites
septatus Vinn, 2005
shell wall in longitudinal section.
© SEM
photographs of cornulitids by Vinn
& Mutvei
Reconstructions of cornulitids
Reconstruction
of Cornulites shell as a lophophorate.